Grahamstoneia humboldti
(agamic)agamic:The agamic (AKA unisexual) generation of an oak gall wasp (cynipini) species consists of only female wasps, which do not mate before laying the eggs which become the male and females of the sexual generation (sexgen).
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The gall's range is computed from the range of all hosts that the gall occurs on. In some cases we have evidence that the gall does not occur across the full range of the hosts and we will remove these places from the range. For undescribed species we will show the expected range based on hosts plus where the galls have been observed.
Our ID Notes may contain important tips necessary for distinguishing this gall
from similar galls and/or important information about the taxonomic status of
this gall inducer.
Created Feb 4, 2026 1:47 PM UTC
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Last updated Feb 4, 2026 1:47 PM UTC
A new genus of Nearctic oak gall wasp, Grahamstoneia Melika & Nicholls, gen. nov. (Hymenoptera: Cynipidae, Cynipini)
George Melika, James Nicholls
(2021)
Grahamstoneia humboldti, sp. nov.
Hosts: Quercus vacciniifolia, chrysolepis
Diagnosis: The gall somewhat resembles that of Andricus projectus Weld, 1952b although is more slender and green when young rather than red. However, morphologically the adults are very different.
Gall. (Fig 12). A bud gall on terminal or axillary buds, up to 8mm long and 2–3mm in diameter. Smooth conical gall with a pointed tip, emerging from bud scales. Young galls are green with a slight pubescence at base that rubs off easily; mature galls are a rich dark brown colour.
Biology. Only the asexual generation is known, which induces galls on Q. vacciniifolia and Q. chrysolepis within Quercus section Protobalanus – the golden cup or intermediate oaks (Denk et al. 2017, Govaerts & Frodin 1998). Mature galls were collected in early November, and adults emerged soon afterwards.
Distribution. USA: California (three locations: Berry summit, ~25km E of Arcata; a site 30km NE of Arnold; and Irvine Lodge rest area on route 101, ~20km N of Willits); Arizona (Oak Creek Canyon, 15km N of Sedona).