The rust fungus Chrysomyxa woroninii causes perennial witches' brooms on several species of Ledum in northern and subalpine regions of Europe, North America and Asia. Spruce bud rust has been assumed to be the aecial state of C. woroninii because of the close proximity of infected Ledum plants and systemically infected buds on Picea. Spruce bud rust results in stunted shoot formation when buds and female cones are infected. The disease also has life cycle stages on Labrador tea.

- Patricia E. CRANE, Yasuyuki HIRATSUKA, Randolph S. CURRAH: (1999) Clarification of the life-cycle of Chrysomyxa woroninii on Ledum and Picea^©