Pachypsylla venusta

Pachypsylla celtidisvesicula

Pachypsylla celtidisasterisca

Pachypsylla celtidisumbilicus

Pachypsylla celtidismamma

Pachypsylla celtidisinteneris

Pachypsylla pallida

Pachypsylla celtidisgemma

Pachypsylla celtidiscucurbita

Pachypsylla celtidisglobula

Pachypsylla celtidispubescens

Pachypsylla laevovesicula

Pachypsylla reticulovesicula

Pachypsylla rugosa

Pachypsylla c-laevigata-blister-gall

Pachypsylla venusta

Pachypsylla celtidisgrandis
Pachypsylla tridentata

Hosts: Celtis occidentalis, tenuifolia, laevigata, reticulata

Galls spherical and large (1-3 cm) on petiole, usually polythalamous

All over the hosts ranges, northeast to CT, southwest to AZ, UT, southeast to Georgia, northwest to ID.

Pachypsylla celtidiscesicula

Celtis occidentalis

Galls inconspicuous, blister like, elevated on both upper and under sides of the leaf

Range: Northeastern US, as for C occidentalis 

Star gall with upper central spine

Pachypsylla celtidisasterisca

Celtis laevigata, reticulata

Galls rounded, raised on the upper side; star-shaped or flower-shaped on the under side

Star gall without upper central spine

Pachypsylla celtidisasterisca

Celtis tenuifolia, (occidentalis) [parentheses indicate occasional observation]

Pachypsylla celtidisumbilicus

Celtis occidentalis

Galls disc-like or button-like, much flattened; upper depression with an outer rim and the central portion slightly raised with a median spine; under side wart-like, with a depression at middle

MD, VA, AR, MO, IA, MS

Pachypsylla celtidismamma

Celtis occidentalis

Hairy nipple galls. Galls pubescent on the apical portion of the under side, but hairs easily rubbed off, giving glabrous appearance; upper side a  depression with central spine but no outer rim; under side nipple-like, longer than wide, without obvious central depression 

Widely distributed in eastern North America, north to Ontario, south to NC, east to CT, NY, west to NE, KS, OK

Pachypsylla rohweri

Celtis reticulata

Hairy nipple galls

CO, TX, OK

Conclusions about the status of several additional described nipple gall makers, including P. celtidispubescens, P. rohweri, P. celtidisglobulus, P. celtidiscucurbita and P. celtidiscucurbita var."?", can not be made, since they were not included in this study. 

Pachypsylla celtidisinteneris

galls oblong-oval in shape on one side of the twig, monothalamous

Pachypsylla pallida

Celtis reticulata

gall densely pubescent on bud,  usually polythalamous

Within the bud gallers, allozymes show little geographic variation, even between allopatric gall morphs, which argues against the separate species names, P. celtidisgemma Riley (glabrous bud gall) and P. pallida Patch (hairy bud gall) advanced for these morphs by some authors. However, the gall morphs are associated with differences in wing pattern and host affiliation. Within the petiole galls, there is significant allozyme differentiation among geographic populations on different hosts. The possibility that allopatric populations of the petiole and bud gallers are specifically distinct remains open.

AZ, NM, OK

Pachypsylla celtidisgemma

Celtis occidentalis, tenuifolia, laevigata

Gall glabrous on bud, usually polythalamous

Within the bud gallers, allozymes show little geographic variation, even between allopatric gall morphs, which argues against the separate species names, P. celtidisgemma Riley (glabrous bud gall) and P. pallida Patch (hairy bud gall) advanced for these morphs by some authors. However, the gall morphs are associated with differences in wing pattern and host affiliation. Within the petiole galls, there is significant allozyme differentiation among geographic populations on different hosts. The possibility that allopatric populations of the petiole and bud gallers are specifically distinct remains open.

Central and eastern US

Pachypsylla celtidiscucurbita [not listed in the key in which Yang provides gall descriptions, included in Table 1-2 listing described species without endorsing their validity or updating info]

Host: Celtis tenuifolia

Gall: Glabrous nipple gall

Range: Northeastern US, eg MD, VA, MO, MS


P. sp. Riemann 1961 (probably=P celtidiscurcurbita var ? Riley 1890)

Celtis laevigata

Gall: Glabrous nipple gall

Southern US, eg MO, TX

Conclusions about the status of several additional described nipple gall makers, including P. celtidispubescens, P. rohweri, P. celtidisglobulus, P. celtidiscucurbita and P. celtidiscucurbita var."?", can not be made, since they were not included in this study. 

Pachypsylla celtidisglobulus

Celtis tenuifolia

Glabrous nipple gall. Galls glabrous; upper side forming a cup-like depression with an outer rim; under side rounded, nearly as long as wide or wider than length, apple-like, with a central depression

Northeastern US, eg MD, VA, MO, MS

Though more evidence is needed, it is probable that in addition to the blister galler, the three species named by Riley but synonymized by others (e.g. Crawford, 1914; Tuthill, 1943) , i.e. P. ,celtidisumbilicus (disc galler), P. celtidisglobulus (glabrous nipple galler) and P. celtidisasterisca (star galler), should be revived. 

Conclusions about the status of several additional described nipple gall makers, including P. celtidispubescens, P. rohweri, P. celtidisglobulus, P. celtidiscucurbita and P. celtidiscucurbita var."?", can not be made, since they were not included in this study. 

Pachypsylla celtidispubescens [not listed in the key in which Yang provides gall descriptions, included in Table 1-2 listing described species without endorsing their validity or updating info]

Celtis reticulata

Southwestern U. S., e.g. TX, OK, CO, MS.

Conclusions about the status of several additional described nipple gall makers, including P. celtidispubescens, P. rohweri, P. celtidisglobulus, P. celtidiscucurbita and P. celtidiscucurbita var."?", can not be made, since they were not included in this study. 

new sp. 1 Riemann 1961

Celtis laevigata

Regular blister gall without upper central spine

Texas

new sp. 2 Riemann 1961

Host: Celtis reticulata

Regular blister gall without upper central spine

Range: TX, NM, AZ, OK

new sp. 3 Riemann 1961

Hosts: Celtis reticulata (laevigata)

Rough blister gall without upper central spine

TX

?

Hosts: Celtis laevigata, (tenuifolia) [parentheses indicate occasional observation]

Blister gall with upper central spine

Range: LA, VA

Biosystematics and the evolution of gall formation in hackberry psyllids Pachypsylla (Insecta: Homoptera: Psylloidea: Psyllidae)