Amphibolips quercusostensackenii (sexgen)

Andricus dimorphus (agamic)

Andricus nigricens (agamic)

Andricus weldi (agamic)

Sphaeroteras pulchripennis (agamic)

Feron clarkei (sexgen)

Amphibolips quercusrugosa (sexgen)

Amphibolips ellipsoidalis

Trigonaspis quercusforticorne (agamic)

Trigonaspis radicola (sexgen)

Phylloteras campanulatum (agamic)

Biorhiza caepuliformis (agamic)

Feron comatum (sexgen)

Amphibolips femoratus (sexgen)

Andricus perfulvum (agamic)

Nichollsiella arizonica (agamic)

Dros sessile (agamic)

Feron amphorus (agamic)

Dros viscidum (agamic)

Trigonaspis fumosa (sexgen)

Trigonaspis mediocre (agamic)

Cynips conspicuus (agamic)

Cynips heldae (agamic)

Amphibolips quercuspomiformis (agamic)

Acraspis gemula (sexgen)

Cynips quercusechinus (sexgen)

Atrusca trimaculosa (agamic)

Cynips quercusechinus (agamic)

A quercusostensackenii, comb. nov. (=Andricus quercussingularis (Bassett, 1863), syn. nov.)

Andricus dimorphus (Beutenmueller), comb. nov.

Andricus nigricens (Gillette), comb. nov.

Andricus weldi (Beutenmueller), comb. nov.

Amphibolips quercusrugosa comb. nov

herein transferred to the Amphibolips genus

Amphibolips ellipsoidalis comb. rev.

6 species from the genus Andricus herein are transferred to Amphibolips: A ellipsoidalis Weld, 1926 comb. rev.

Trigonaspis quercusforticorne (Weld) comb. nov.

Seven Xanthoteras species here are transferred to Trigonaspis. . . Trigonaspis quercusforticorne (Weld) comb. nov.


Trigonaspis radicola (Ashmead) comb. nov.

Seven Xanthoteras species here are transferred to Trigonaspis. . . Trigonaspis radicola (Ashmead) comb. nov.


Phylloteras campanulatum comb. nov.

Lyon (1993) separated Euxystoteras from Phylloteras "by its simple tarsal claws". The  inadequacy  of  this  generic  character  argues  that  Euxystoteras  must  be  a  synonym  of  Phylloteras.  Thus,  Euxystoteras  campanulatum  (Lyon  1993)  must  be  transferred  to  Phylloteras, P. campanulatum (Lyon), comb. nov.

Biorhiza caepuliformis comb. rev.

[moves caepuliformis back from Sphaeroteras to Biorhiza]

Andricus comata

Consequently,  three  Liodora species  known  from  North  America  must  be  transferred  to  Andricus: comata (Weld), comb.  nov.

Amphibolips femoratus

On the basis of diagnostic characters given above, 6 species from the genus Andricus herein  are  transferred  to  Amphibolips: A. femoratus Ashmead,  1887,  comb. nov.

Consequently, all 8 known species [in the genus Trichoteras] must be transferred to Andricus: perfulvum (Weld), comb. nov.

One  species,  [Adleria] arizonica  (Cockerell)  must  be  transferred  to  Disholcaspis: D. arizonicus (Cockerell), comb. nov

transferred to Andricus: sessilum (Weld), comb. nov.

transferred  to  Andricus: amphorus  (Weld),  comb.  nov.

transferred  to  Andricus: viscidus (Weld), comb. nov.

transferred  to  Trigonaspis: fumosa  Weld,  comb.  rev

 transferred  to  Trigonaspis: mediocre  (Weld),  comb.  nov.

The following species must be transferred from Besbicus to Cynips: conspicuus Kinsey, comb.rev.

The following species must be transferred from Besbicus to Cynips: heldae  Fullaway,  comb.  rev.

Two Callirhytis species, quercuspomiformis (Bassett) and quercusrugosa (Ashmead), herein are transferred to the Amphibolips genus.

Acrapis gemula, syn. nov.

Philonix pallipes

Comments. Kinsey (1930) transferred Dryophanta pallipes Bassett into Philonix and considered it as the sexual generation of P. fulvicollis. However, Kinsey provided no justification for  this  synonymy [Kinsey seems to have assigned pallipes to fulvicollis based on a process of elimination]. Weld (1959) wrote that P.  pallipes  was  perhaps  a  synonym  of  Acraspis gemula (Bassett)î. We compared the three specimens of P. pallipes (from Beutenmueller collection, USNM, Washington, DC) with the paratypes of the sexual generation of A. gemula and found no differences. Philonix pallipes is a syn. nov. of A. gemula. If P. pallipes (Bass.) is treated as  an  Acraspis species, there is no evidence that species of Philonix have alternate  sexual generations.

Andricus ribes Kinsey was listed as the sexual generation of A. quercusechinus (Weld 1951; Burks 1979). Weld (1952a) described for this species that "on circumstantial evidence this is considered the alternating generation of echinus." There is no evidence for this. Kinsey (1922) did not mention that A. ribes might be the sexual generation of any other cynipid wasp in the original description. Kinsey (1930) gave no acceptable evidence for the synonymization of A. ribes to A. quercusechinus. However, A. ribes is entirely congeneric with sexual Cynips and, of course, is the sexual generation of one of the Californian Cynips (Antron) species. However, it is too early to state that it is the sexual generation of A. quercusechinus; this must be proved. 

The following species must be transferred to Cynips: as well as the following Antron species: comb. rev., quercusechinus Osten Sacken

Atrusca trimaculosa (McCracken and Egbert), comb. nov

five species from the Sphaeroteras genus herein are transferred into Atrusca


The following species must be transferred to Cynips: as well as the following Antron species: comb. rev., quercusechinus Osten Sacken

Review of the world genera of oak cynipid wasps (Hymenoptera: Cynipidae: Cynipini)